Dr. Greenberg graduated from Washington University with a degree in biology. He received his M.D. summa cum laude from the State University of New York, Downstate Medical Center, in 1971. After completing postdoctoral training at the University of California at San Diego, he joined the Fred Hutchinson Cancer Research Center and the Division of Oncology at the University of Washington in 1976. He is currently the Head of the Program in Immunology at the Fred Hutchinson Cancer Research Center.

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Full Member, Appointed: 1999, Fred Hutchinson Cancer Research Center, Clinical Research Division, Program in Immunology; Professor, Appointed: 2000, Department of Medicine, Division of Medical Oncology;  and Adjunct Professor, Appointed 2015: Department of Immunology, University of Washington, School of Medicine

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